amf-phylogeny

Glomeromycota SPECIES LIST by Arthur Schüßler and Chris Walker

last updated Januar 17, 2012 - News: We will soon try to update this list. A large number of taxa have been described by the group of Fritz Oehl and coworkers, but some are monospecific or based on doubtful assumptions. Clearly some of these taxa will stay as they will have proirity, according to the Botanical Code, but others are synonyms, or based on wrong sequence information (e.g. of two Scutellospora species). We will try to screen the options on how to show these data, here.
A new Racocetra and one "Cetraspora" (a genus rejected by Morton and Msiska 2010) species were added. Sclerocystis was published in 1873, not 1875, this is corrected.

The Gigasporaceae systematics was adopted according to Morton and Msiska (2010), except that Racocetra weresubiae is treated sensu Oehl et al. (2008), because of its molecular phylogenetic placement. For obious and long known reasons we follow the rejection of Kuklospora, which members now have been placed in Acaulospora (Kaonongbua et al. 2010). Intraspora was synonymised with Archaeospora, a number of species was transferred from Glomus to Diversispora, and the Glomeraceae were split in two families, after the placement of the generic type of Glomus, Glomus macrocarpum, could be resolved (Schüßler and Walker 2010). This species list of the Glomeromycota published in several international libraries is also free availabe as an online copy (Schüßler and Walker 2010) and as a printed copy, including a 'corrigendum'. The aim of our work is a stable, molecular phylogeny based systematics of the Glomeromycota.

If you spot mistakes, please inform us - we try to hold everything up to date and collaborate with the Index Fungorum and Species 2000 databases.

Thanks to those which already sent us pdf-files, or scanned pages, and to the publishers that gave us copyright clearance (see list at the end of the table).

We have been refused clearance by Springer and by the journals Nova Hedwigia and Botany (former Can. J. Bot.), and thus we cannot provide pdf files of the respective papers. We suggest that authors choose journals with suitable policy, or you may prepare an 'authors version' (pre-publisher layout) that usually is allowed to be freely placed online. We are happy to link to such papers hosted at your institutional servers (if you provide us with the hyperlink).

Go directly to the genera (sorted alphabetically, in the table below):

Paraglomerales: Paraglomus
Archaeosporales: Ambispora   Archaeospora   Geosiphon
Diversisporales: Acaulospora   Diversispora   Entrophospora   Gigaspora   Otospora   Pacispora   Racocetra   Redeckera   Scutellospora
Glomerales: Claroideoglomus   Funneliformis   Glomus   Rhizophagus   Sclerocystis

Colour coding in the following table: taxon in blue = hyperlinked to description (pdf-file); green = additional information or opinions.

For references see Schüßler A, Walker C (2010) The Glomeromycota: a species list with new families and new genera. Schüßler A, Walker C, Gloucester; published in libraries at Royal Botanic Garden Edinburgh Kew, Botanische Staatssammlung Munich, and Oregon State University. This species list of the Glomeromycota published in several international libraries is also free availabe as an online copy (Schüßler and Walker 2010) and as a printed copy, including a 'corrigendum'.

*Glomeromycota*
Current name	Basionyms, synonyms & additional comments	Authorities	Family	Order
*Acaulospora*	back to top	Trappe & Gerd. (1974)	Acaulosporaceae	Diversisporales
*Acaulospora alpina*	Classification based on molecular and morphological evidence from field material.	Oehl, Sykorova & Sieverd. (2006)	Acaulosporaceae	Diversisporales
*Acaulospora cavernata*	A culture, at first considered to be Acaulospora scrobiculata, was established from the UK, and registered as BEG33. The culture was established from five spores that all appeared to have ornamentation similar to A. scrobiculata but the spores produced now are predominantly similar to the type of A. cavernata. The type material of A. cavernata is not suitable for molecular study. BEG33 was designated as an epitype for the species.	Błaszk. (1989)	Acaulosporaceae	Diversisporales
*Acaulospora colliculosa*	Species not available in culture; the molecular analysis was made from authenticated specimens taken from the field. It is closely related and als similar in spore morphology to A. brasiliensis.	Kaonongbua, J.B. Morton & Bever (2010)	Acaulosporaceae	Diversisporales
*Acaulospora colombiana*	Basionym: Entrophospora colombiana Spain & N.C. Schenck (1984) Synonyms: Kuklospora colombiana Oehl & Sieverd. (2006)	(Spain & N.C. Schenck) Kaonongbua, J.B. Morton & Bever (2010)	Acaulosporaceae	Diversisporales
*Acaulospora delicata*		C. Walker, C.M. Pfeiff. & Bloss (1986)	Acaulosporaceae	Diversisporales
*Acaulospora dilatata*		J.B. Morton (1986)	Acaulosporaceae	Diversisporales
*Acaulospora entreriana*		M.S. Velázquez & Cabello (2008)	Acaulosporaceae	Diversisporales
*Acaulospora foveata*	Molecular evidence from two INVAM cultures, CR315 and CR401A. The species description does not illustrate the full amount of variation seen in the spore ornamentation of the type material. The ornamentation illustrated for culture BR861 on the INVAM website is not entirely consistent with that of the type material for the species. It seems likely that spore ornamentation is somewhat variable.	Trappe & Janos (1982)	Acaulosporaceae	Diversisporales
*Acaulospora kentinensis*	Basionym: Entrophospora kentinensis C.G. Wu & Y.S. Liu (1995) Synonyms: Kuklospora kentinensis Oehl & Sieverd. (2006)	(C.G. Wu & Y.S. Liu) Kaonongbua, J.B. Morton & Bever (2010)	Acaulosporaceae	Diversisporales
*Acaulospora koskei*		Błaszk. (1995)	Acaulosporaceae	Diversisporales
*Acaulospora lacunosa*		J.B. Morton (1986)	Acaulosporaceae	Diversisporales
*Acaulospora laevis*		Gerd. & Trappe (1974)	Acaulosporaceae	Diversisporales
*Acaulospora longula*		Spain & N.C. Schenck (1984)	Acaulosporaceae	Diversisporales
*Acaulospora mellea*		Spain & N.C. Schenck (1984)	Acaulosporaceae	Diversisporales
*Acaulospora morrowiae*		Spain & N.C. Schenck (1984) [as 'morrowae']	Acaulosporaceae	Diversisporales
*Acaulospora paulinae*	Compare with A. undulata. Species descriptions of A. undulata and A. paulinae are very similar. One of the differences that was considered important in separating the species was that A. undulata did not react in Melzer’s reagent. However, this reaction in members of Acaulospora varies with condition of the specimens, and Kaonongbua et al (2010) (see A. colliculosa) have indicated that reaction to this reagent in any case may not be a useful phylogenetic signal.	Błaszk. (1988)	Acaulosporaceae	Diversisporales
*Acaulospora scrobiculata*		Trappe (1977)	Acaulosporaceae	Diversisporales
*Acaulospora spinosa*		C. Walker & Trappe (1981)	Acaulosporaceae	Diversisporales
*Acaulospora tuberculata*		Janos & Trappe (1982)	Acaulosporaceae	Diversisporales
*Acaulospora*	species of uncertain position
*Acaulospora* *bireticulata*	Probably a synonym for A. elegans, which can be ‚transformed‘ into A. bireticulata by use of Nomarski differential interference microscopy; it is likely that this species is a heterotypic synonym of the former.	F.M. Rothwell & Trappe (1979)
*Acaulospora capsicula*	Has many similarities with A. laevis, but is described as having a reaction to Melzer’s reagent.	Błaszk. (1990)
*Acaulospora colossica*	Is similar to A. laevis. It is described as lacking a reaction to Melzer’s reagent.	P.A. Schultz, Bever & J.B. Morton (1999)
*Acaulospora denticulata*		Sieverd. & S. Toro (1987)
*Acaulospora elegans*	Compare with A. bireticulata	Trappe & Gerd. (1974)
*Acaulospora excavata*	The ornamentation on this species is of large deep cavities, leading to the suspicion that it belongs with such organisms as A. scrobiculata.	Ingleby & C. Walker (1994)
*Acaulospora gedanensis*		Błaszk. (1988)
*Acaulospora myriocarpa*		Spain, Sieverd. & N.C. Schenck (1986)
*Acaulospora nicolsonii*	This species probably should be in Ambispora, but must remain in Acaulospora pending molecular or other evidence.	C. Walker, L.E. Reed & F.E. Sanders (1984)
*Acaulospora polonica*		Błaszk. (1988)
*Acaulospora rehmii*		Sieverd. & S. Toro (1987)
*Acaulospora rugosa*		J.B. Morton (1986)
*Acaulospora splendida*	This species produces acaulosporoid spores, but they do not seem to share characteristics with other members of the genus. As there is no molecular data available the species is retained in its current genus.	Sieverd., Chaverri & I. Rojas (1988)
*Acaulospora sporocarpia*		S.M. Berch (1985)
*Acaulospora taiwania*	Very unlikely to be an Acaulospora sp. In the species description, it is described as being produced in sporocarps. The illustrations of the species are of such poor quality that it is impossible to draw conclusions other than that it does not look much like an acaulospore producing fungus. The species description itself is not much more help, and the type material is lost.	H.T. Hu (1988)
*Acaulospora thomii*	Is said to be fundamentally different from other species in the genus, and it has been considered as a possible member of Archaeospora. One major consideration was the lack of reaction to Melzer’s reagent, but this is not unknown among the ornamented, small-spored species in Acaulospora. Until more is known recombining it within a different genus would be speculative.	Błaszk. (1988)
*Acaulospora undulata*	Compare with A. paulinae, see comments given for A. paulinae.	Sieverd. (1988)
*Acaulospora walkeri*	Known only from field collected material, this species is close in morphology to other species in the genus with small, smooth, yellow spores.	Kramad. & Hedger (1990)
*Ambispora*	back to top	C. Walker, Vestberg & Schuessler (2007)	Ambisporaceae	Archaeosporales
*Ambispora appendicula*	Basionym: Acaulospora appendicula Spain, Sieverd. & N.C. Schenck (1984) Synonyms: Appendicispora appendicula Spain, Oehl & Sieverd. (2006), Paracaulospora appendicula S.P. Gautam & U.S. Patel (2007)	(Spain, Sieverd. & N.C. Schenck) C. Walker (2008)	Ambisporaceae	Archaeosporales
*Ambispora brasiliensis*		B.T. Goto, L.C. Maia & Oehl (2008)	Ambisporaceae	Archaeosporales
*Ambispora callosa*	Basionym: Glomus callosum Sieverd. (1988) Synonyms: Appendicispora callosa C. Walker, Vestberg & Schuessler (2007)	(Sieverd.) C. Walker, Vestberg & Schuessler (2007)	Ambisporaceae	Archaeosporales
*Ambispora fecundispora*	Basionym: Glomus fecundisporum N.C. Schenck & G.S. Sm. (1982) Synonyms: Appendicispora fecundispora C. Walker, Vestberg & Schuessler (2007)	(N.C. Schenck & G.S. Sm.) C. Walker (2008)	Ambisporaceae	Archaeosporales
*Ambispora fennica*	Synonyms: Appendicispora fennica C. Walker, Vestberg & Schuessler (2007)	C. Walker, Vestberg & Schuessler (2007)	Ambisporaceae	Archaeosporales
*Ambispora gerdemannii*	Basionym: Glomus gerdemannii S.L. Rose, B.A. Daniels & Trappe (1979) Synonyms: Appendicispora gerdemannii Spain, Oehl & Sieverd. (2006), Archaeospora gerdemannii J.B. Morton & D. Redecker (2001)	(S.L. Rose, B.A. Daniels & Trappe) C. Walker, Vestberg & Schuessler (2007)	Ambisporaceae	Archaeosporales
*Ambispora granatensis*		J. Palenzuela, N. Ferrol & Oehl (2011)	Ambisporaceae	Archaeosporales
*Ambispora jimgerdemannii*	Basionym: Acaulospora gerdemannii N.C. Schenck & T.H. Nicolson (1979) Synonyms: Appendicispora jimgerdemannii Spain, Oehl & Sieverd. (2006)	(N.C. Schenck & T.H. Nicolson) C. Walker (2008)	Ambisporaceae	Archaeosporales
*Ambispora leptoticha*	Basionym: Glomus leptotichum N.C. Schenck & G.S. Sm. (1982) Synonyms: Appendicispora leptoticha C. Walker, Vestberg & Schuessler (2007), Archaeospora leptoticha J.B. Morton & D. Redecker (2001), Pseudoglomus leptotichum S.P. Gautam & U.S. Patel (2007) Remark: Archaeospora leptoticha J.B. Morton & D. Redecker (2001) was composed of Ac. gerdemannii & G. leptotichum, Ac. appendicula & G. fecundisporum (Morton et al. 1997). This was eventually revised by Spain et al. (2006) and Walker (2008). For a full explanation about what happened to the species Acaulospora gerdemannii, Ac. appendicula, Glomus leptotichum and Gl. fecundisporum, see here.	(N.C. Schenck & G.S. Sm.) C. Walker, Vestberg & Schuessler (2007)	Ambisporaceae	Archaeosporales
*Archaeospora*	back to top	J.B. Morton & D. Redecker (2001)	Archaeosporaceae	Archaeosporales
*Archaeospora schenckii*	Basionym: Entrophospora schenckii Sieverd. & S. Toro (1987) Synonyms: Intraspora schenckii Oehl & Sieverd. (2006)	(Sieverd. & S. Toro) Walker & Schuessler (2010)	Archaeosporaceae	Archaeosporales
*Archaeospora trappei* (emendation by Spain 2003)	Basionym: Acaulospora trappei R.N. Ames & Linderman (1976)	(R.N. Ames & Linderman) J.B. Morton & D. Redecker (2001)	Archaeosporaceae	Archaeosporales
*Claroideoglomus*	back to top	C. Walker & Schuessler (2010)	Claroideoglomeraceae	Glomerales (emendation)
*Claroideoglomus claroideum*	Basionym: Glomus claroideum N. C. Schenck & G. S. Sm. (1982) (emendation by Walker & Vestberg 1998) Synonyms: Glomus maculosum D. D. Mill. & C. Walker (1986), G. fistulosum Skou & I. Jakobsen (1989)	(N. C. Schenck & G. S. Sm.) C. Walker & Schuessler (2010)	Claroideoglomeraceae	Glomerales
*Claroideoglomus drummondii*	Basionym: Glomus drummondii Blaszk. & C. Renker (2006)	(Blaszk. & C. Renker) C. Walker & Schuessler (2010)	Claroideoglomeraceae	Glomerales
*Claroideoglomus etunicatum*	Basionym: Glomus etunicatum W.N. Becker & Gerd. (1977) [as 'etunicatus']	(W.N. Becker & Gerd.) C. Walker & Schuessler (2010)	Claroideoglomeraceae	Glomerales
*Claroideoglomus lamellosum*	Basionym: Glomus lamellosum Dalpé, Koske & Tews (1992)	(Dalpé, Koske & Tews) C. Walker & Schuessler (2010)	Claroideoglomeraceae	Glomerales
*Claroideoglomus luteum*	Basionym: Glomus luteum L.J. Kenn., J.C. Stutz & J.B. Morton (1999)	(L.J. Kenn., J.C. Stutz & J.B. Morton) C. Walker & Schuessler (2010)	Claroideoglomeraceae	Glomerales
*Claroideoglomus walkeri*	Basionym: Glomus walkeri Blaszk. & C. Renker (2006)	(Blaszk. & C. Renker) C. Walker & Schuessler (2010)	Claroideoglomeraceae	Glomerales
*Diversispora*	back to top	C. Walker & Schuessler (2004)	Diversisporaceae	Diversisporales
*Diversispora aurantia*	Basionym: Glomus aurantium Błaszk., Blanke, Renker & Buscot (2004)	(Błaszk., Blanke, Renker & Buscot) C. Walker & Schuessler (2010) [as 'aurantium']	Diversisporaceae	Diversisporales
*Diversispora celata*		C. Walker, Gamper & Schuessler (2009)	Diversisporaceae	Diversisporales
*Diversispora eburnea*	Basionym: Glomus eburneum L.J. Kenn., J.C. Stutz & J.B. Morton (1999)	(L.J. Kenn., J.C. Stutz & J.B. Morton) C. Walker & Schuessler (2010)	Diversisporaceae	Diversisporales
*Diversispora epigaea*	Basionym: Glomus epigaeum B.A. Daniels & Trappe (1979)	(B.A. Daniels & Trappe) C. Walker & Schuessler (2010)	Diversisporaceae	Diversisporales
*Diversispora trimurales*	Basionym: Glomus trimurales Koske & Halvorson (1989)	(Koske & Halvorson) C. Walker & Schuessler (2010)	Diversisporaceae	Diversisporales
*Diversispora spurca* (emendation)	Basionym: Glomus spurcum C.M. Pfeiff., C. Walker & Bloss (1996)	(C.M. Pfeiff., C. Walker & Bloss) C. Walker & A. Schuessler (2004) [as 'spurcum']	Diversisporaceae	Diversisporales
*Entrophospora*	back to top Molecular data for E. infrequens do not provide evidence of its phylogeny. Morphologically, there seems little reason to maintain the genus, but clarification must await evidence. Molecular phylogenetic and systematic position of Entrophosporaceae therefore unknown.	R.N. Ames & R.W. Schneid. (1979)	Entrophosporaceae
*Entrophospora infrequens*	Basionym: Glomus infrequens I.R. Hall (1977)	(I.R. Hall) R.N. Ames & R.W. Schneid. (1979)	Entrophosporaceae
*Entrophospora*	species of uncertain position
*Entrophospora baltica*		Błaszk., Madej & Tadych (1998)
*Entrophospora nevadensis*	rDNA sequences published cluster in Diversispora	J. Palenzuela, N. Ferrol, Azcón-Aguilar & Oehl (2010)	sequences cluster in Diversisporaceae	sequences cluster in Diversisporales
*Funneliformis*	back to top	C. Walker & Schuessler (2010)	Glomeraceae Pirozynsky & Dalpé (1989)	Glomerales (emendation)
*Funneliformis africanum*	Basionym: Glomus africanum Błaszk. & Kovács (2010)	(Błaszk. & Kovács) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Funneliformis badius*	Basionym: Glomus badium Oehl, Redecker & Sieverd. (2005)	(Oehl, Redecker & Sieverd.) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Funneliformis caledonius*	Basionym: Endogone macrocarpa var. caledonia T.H. Nicolson & Gerd. (1968) Synonyms: Glomus caledonium Trappe & Gerd. [as 'caledonius'] (1974)	(T.H. Nicolson & Gerd.) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Funneliformis coronatus*	Basionym: Glomus coronatum Giovann. (1991)	(Giovann.) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Funneliformis mosseae*	Basionym: Endogone mosseae T.H. Nicolson & Gerd. (1968) Synonyms: Glomus mosseae Gerd. & Trappe (1974)	(T.H. Nicolson & Gerd.) C. Walker & Schuessler 2010	Glomeraceae	Glomerales
*Funneliformis fragilistratus*	Basionym: Glomus fragilistratum Skou & I. Jakobsen (1989)	(Skou & I. Jakobsen) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Funneliformis geosporus*	Basionym: Endogone macrocarpa var. geospora T.H. Nicolson & Gerd. (1968) Synonyms: Glomus macrocarpum var. geosporum Gerd. & Trappe (1974), Glomus geosporum C. Walker (1982) (complete paper here)	(T.H. Nicolson & Gerd.) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Funneliformis verruculosus*	Basionym: Glomus verruculosum Błaszk. (1997)	(Błaszk.) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
Funneliformis vesiculiferus NOTE: this is a Rhizophagus species.	Basionym: Endogone vesiculifera Thaxt. (1922) Synonyms: Glomus vesiculiferum Gerd. & Trappe (1974) [as 'vesiculifer'] Formally the species was erroneously placed in Funneliformis in Schüßler & Walker, 2010; this will be corrected soon.	(Thaxt.) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Funneliformis constrictus*	Basionym: Glomus constrictum Trappe (1977) [as 'constrictus']	(Trappe) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Funneliformis xanthium*	Basionym: Glomus xanthium Błaszk., Blanke, Renker & Buscot (2004)	(Błaszk., Blanke, Renker & Buscot) C. Walker & Schuessler (2010)	Glomeraceae	Glomerales
*Geosiphon*	back to top	F. v. Wettstein (1915)	Geosiphonaceae (emendation)	Archaeosporales
*Geosiphon pyriformis* (emendation)	Basionym: Botrydium pyriforme Kützing (1849) Synonyms: Geosiphonomyces pyriformis Cif. & Tomas (1957)	(Kützing) F. v. Wettstein (1915) [as 'pyriforme']	Geosiphonaceae	Archaeosporales
*Gigaspora*	back to top	Gerd. & Trappe (1974)	Gigasporaceae	Diversisporales
*Gigaspora albida*		N.C. Schenck & G.S. Sm. (1982)	Gigasporaceae	Diversisporales
*Gigaspora candida*	Synonyms: Gigaspora alboaurantiaca W.N. Chou (1991) The type material of G. candida is lost, but the species description is such that it cannot be separated from G. alboaurantiaca. Therefore, G. alboaurantiaca was placed as a later heterotypic synonym of G. candida.	Bhattacharjee, Mukerji, J.P. Tewari & Skoropad (1982)	Gigasporaceae	Diversisporales
*Gigaspora decipiens*		I.R. Hall & L.K. Abbott (1984)	Gigasporaceae	Diversisporales
*Gigaspora gigantea*	Basionym: Endogone gigantea T.H. Nicolson & Gerd. (1968)	(T.H. Nicolson & Gerd.) Gerd. & Trappe (1974)	Gigasporaceae	Diversisporales
*Gigaspora margarita*		W.N. Becker & I.R. Hall (1976)	Gigasporaceae	Diversisporales
*Gigaspora ramisporophora*		Spain, Sieverd. & N.C. Schenck (1989)	Gigasporaceae	Diversisporales
*Gigaspora rosea*		T.H. Nicolson & N.C. Schenck (1979)	Gigasporaceae	Diversisporales
*Gigaspora*	species of uncertain position
*Gigaspora lazzarii*	Type material and species description share none of the characteristics of any member of the Glomeromycota.	Montecchi, Ruini & G. Gross (1996)
*Glomus* sensu stricto	back to top Synonym: Parapseudoglomus S.P. Gautam & U.S. Patel (2007)	Tul. & C. Tul. (1845)	Glomeraceae Pirozynsky & Dalpé (1989)	Glomerales (emendation)
*Glomus macrocarpum* (lectotypified by Berch & Fortin 1983)	Basionym: Glomus macrocarpum Tul. & C. Tul. (1845) Synonyms: Endogone macrocarpa Tul. & C. Tul. (1851), Glomus macrocarpum var. macrocarpum	Tul. & C. Tul. (1845) [as 'macrocarpus']	Glomeraceae	Glomerales
*Glomus* sensu lato	species of uncertain position
*Glomus achrum*	According to analysis of published rDNA sequence data, this species clusters basal in the Glomeraceae in the genus Rhizophagus, but the phylogenetic position cannot yet be robustly defined.	Błaszk., D. Redecker, Koegel, Schuezek, Oehl & Kovacs (2009)	Glomeraceae	Glomerales
*Glomus aggregatum* (emendation here)	According to published β-tubulin sequence data, G. aggregatum clusters with Claroideoglomus, but this seems unlikely, so it is classified as of uncertain position, for now	N.C. Schenck & G.S. Sm. (1982)
*Glomus albidum*	Morphologically, the species seems identical to D. eburneum	C. Walker & L.H. Rhodes (1981)
*Glomus ambisporum*		G.S. Sm. & N.C. Schenck (1985)
*Glomus antarcticum*	Morphologically similar to Rhizophagus species	Cabello (1994)
*Glomus arborense*		McGee (1986)
*Glomus arenarium*		Błaszk., Tadych & Madej (2001)
*Glomus atrouva*	Compare with Glomus botryoides	McGee & Pattinson (2002)
*Glomus aureum*	Compare with Glomus invermaium	Oehl & Sieverd. (2003)
*Glomus australe*	Basionym: Endogone australis Berk. (1859) [1860]	(Berk.) S.M. Berch (1983)
*Glomus avelingiae*		R.C. Sinclair (2000)
*Glomus bagyarajii*		V.S. Mehrotra (1997)
*Glomus bistratum*	According to analysis of published rDNA sequences, the phylogenetic position cannot yet be robustly defined	Błaszk., D. Redecker, Koegel, Symanczik, Oehl & Kovacs (2009)	Glomeraceae	Glomerales
*Glomus boreale*	Basionym: Endogone borealis Thaxt. (1922)	(Thaxt.) Trappe & Gerd. (1974)
*Glomus botryoides*		F.M. Rothwell & Victor (1984)
*Glomus caesaris*		Sieverd. & Oehl (2002)
*Glomus canadense*	Basionym: Endogone canadensis Thaxt. (1922)	(Thaxt.) Trappe & Gerd. (1974)
*Glomus candidum*		Furrazola, Kaonongbua & Bever (2010)
*Glomus canum*		McGee (2002)
*Glomus cerebriforme*	According to published rDNA sequence data, this species clusters in Rhizophagus, but the culture used to generate the sequences may not correspond to the species and is in need of verification, so it is classified as of uncertain position, for now. Compare with Glomus pallidum	McGee (1986)
*Glomus citricola*	Type material difficult to interpret. Maybe mixed, maybe a Rhizophagus sp., or even an Acaulospora	D.Z. Tang & M. Zang (1984) [as 'citricolum']
*Glomus convolutum*		Gerd. & Trappe (1974) [as 'convolutus']
*Glomus corymbiforme*		Błaszk. (1995)
*Glomus cuneatum*		McGee & Cooper (2002)
*Glomus delhiense*		Mukerji, Bhattacharjee & J.P. Tewari (1983)
*Glomus deserticola*	The article in which the molecular evidence for this species is published (Chellappan et al. 2005) shows an illustration of a fungus, but it cannot be verified as G. deserticola. The ex-type culture of this species is available, and pending new molecular evidence, we retain this as a species of uncertain phylogenetic position	Trappe, Bloss & J.A. Menge (1984)
*Glomus dimorphicum*	Confused name due to error. The type includes four already described species	Boyetchko & J.P. Tewari (1986)
*Glomus dolichosporum*		M.Q. Zhang & You S. Wang (1997)
*Glomus flavisporum*	Basionym: Endogone flavispora M. Lange & E.M. Lund (1955)	(M. Lange & E.M. Lund) Trappe & Gerd. (1974) [as ‘flavisporus‘]
*Glomus formosanum*		C.G. Wu & Z.C. Chen (1986)
*Glomus fragile*	Basionym: Paurocotylis fragilis Berk. & Broome (1875)	(Berk. & Broome) Trappe & Gerd. (1974) [as 'fragilis']
*Glomus fuegianum*	Basionym: Endogone fuegiana Speg. (1887)	(Speg.) Trappe & Gerd. (1974) [as 'fuegianus']
*Glomus gibbosum*		Błaszk. (1997)
*Glomus globiferum*		Koske & C. Walker (1986)
*Glomus glomerulatum*		Sieverd. (1987)
Glomus goaensis	poor description, unknown affiliation	Khade (2009)
*Glomus halonatum*	Compare Glomus clarum	S.L. Rose & Trappe (1980) [as 'halonatus']
*Glomus heterosporum*		G.S. Sm. & N.C. Schenck (1985)
*Glomus hoi*	Current DNA evidence for organisms that have been determined as G. hoi shows that two different cultures fall into two clades separated at the level at least of genus and possibly family. Consequently, we are retaining it in the species of uncertain position pending further clarification	S.M. Berch & Trappe (1985)
*Glomus hyderabadensis*		Swarapu, Kunwar, Prasad & Manohar (2004)
*Glomus indicum*	According to published rDNA sequence data, this species clusters basal in the Glomeraceae and probably represents a separate genus that cannot yet be robustly defined	Błaszk., Wubet, Harikumar (2010)	Glomeraceae	Glomerales
*Glomus insculptum*		Błaszk. (2004)
*Glomus invermaium*		I.R. Hall (1977) [as 'invermaius']
*Glomus iranicum*	Synonyms: Rhizophagus iranicus C. Walker & Schuessler (2010) The placement of this species in Rhizophagus (as published in Schüßler and Walker 2010) is not correct. We informally correct thus here, pending formal published correction. This species is phylogenetically not included in Rhizophagus, but is a Glomus species of uncertain affiliation. According to published rDNA sequence data, this species clusters basal in the Glomeraceae, perhaps with G. iranicum, and probably represents a separate genus that cannot yet be robustly defined.	Błaszk., Kovács & Balázs (2010)	Glomeraceae	Glomerales
*Glomus kerguelense*		Dalpé & Strullu (2002)
*Glomus lacteum*	Probably not Glomus	S.L. Rose & Trappe (1980) [as 'lacteus']
*Glomus magnicaul*e		I.R. Hall (1977) [as 'magnicaulis']
*Glomus melanosporum*		Gerd. & Trappe (1974) [as 'melanosporus']
*Glomus microaggregatum*		Koske, Gemma & P.D. Olexia (1986)
*Glomus microcarpum* (lectotypification by S.M. Berch & J.A. Fortin 1984)	Synonyms: Endogone microcarpus Tul. & C. Tul. (1851), E. neglecta	Tul. & C. Tul. (1845) [as 'microcarpus']
*Glomus minutum*		Błaszk., Tadych & Madej (2000) [as 'monosporus']
*Glomus monosporum*	This species was established in pot culture, but the culture was lost long before DNA sequencing methods were applied as markers. There have been several cultured organisms that have been given this name, but none seems properly to fit the species description of brown spores that have ornamentation of spines on the laminated wall component.	Gerd. & Trappe (1974)
*Glomus mortonii*	Although this fungus has produced spores in soil traps in Finland, it has not yet been possible either to establish it in pure culture or to extract DNA. Consequently, it is unclear as to whether or not it belongs in any of the established clades.	Bentiv. & Hetrick (1991)
*Glomus multicaule*		Gerd. & B.K. Bakshi (1976) [as 'multicaulis']
*Glomus multiforum*		Tadych & Błaszk. (1997)
*Glomus* *multisubstensum*	This name has at some time been synonymised with Claroideoglomus claroideum (as Glomus claroideum), but this cannot yet be verified from molecular evidence, and thus has now been moved into uncertain positions	Mukerji, Bhattacharjee & J.P. Tewari (1983)
*Glomus nanolumen*		Koske & Gemma (1990)
*Glomus pallidum*		I.R. Hall (1977) [as 'pallidus']
*Glomus pansihalos*		S.M. Berch & Koske (1986)
*Glomus pellucidum*		McGee & Pattinson (2002)
*Glomus perpusillum*		Błaszk. & Kovács (2009)
*Glomus przelewicense*		Błaszk. (1988) [as 'przelewicensis']
*Glomus pustulatum*		Koske, Friese, C. Walker & Dalpé (1986)
*Glomus radiatum*	Basionym: Endogone radiata Thaxt. (1922)	(Thaxt.) Trappe & Gerd. (1974)
*Glomus reticulatum*	This has such a vague and inadequately illustrated species description that it is impossible to apply the name with any confidence to any organism. The type material seems not to exist, and it is thus a species name that cannot be usefully applied.	Bhattacharjee & Mukerji (1980) [as 'reticulatus']
*Glomus segmentatum*	This is a truly sporocarpic fungus. It is known only from field collections, but no DNA sequence exists that can be used to establish its true phylogeny.	Trappe, Spooner & Ivory (1979) [as 'segmentatus']
*Glomus spinosum*	The species description is more or less uninterpretable, and no type material exists for any of Hu’s species.	H.T. Hu (2002)
*Glomus spinuliferum*	Compare with Glomus pansihalos	Sieverd. & Oehl (2003)
Glomus sterilum this is a nom. inval. (invalid name)	Published without a Latin description or diagnosis (a requirement of the Botanical Code for formal descriptions after 1 Jan 1935).	V.S. Mehrotra & Baijal (1992) [as 'sterile']	none (invalidly published)	none (invalidly published)
*Glomus tenebrosum*	Basionym: Endogone tenebrosa Thaxt. (1922)	(Thaxt.) S.M. Berch (1983)
*Glomus tenerum* (emendation)	A controversial species.	P.A. Tandy (1975) [as 'tener']
*Glomus tenue*	Basionym: Rhizophagus tenuis Greenall (1963) A fungus was published under the name Rhizophagus tenuis and later transferred to the genus Glomus. As used, the name is unlikely to refer to either a single fungus, or a member of Glomus. Fungi fitting the description have from time to time been established in pot culture, but it seems none has ever been purified and there is to date no molecular data with which to place it in its correct clade. It has almost no similarities with any other fungus in the phylum, even more so now that it is known that glomoid spores have been shown to occur in many widely separated taxa.	(Greenall) I.R. Hall (1977) [as 'tenuis']
*Glomus tortuosum*		N.C. Schenck & G.S. Sm. (1982)
*Glomus versiforme*	Basionym: Endogone versiformis P. Karst (1884), non sensu Berch & Fortin, Can. J. Bot. 61: 2614 (1983). This species was confounded by the erroneous synonymisation of Glomus epigaeum with G. versiforme. These two species have been shown not to be synonymous, but the placing of G. versiforme is unknown..	(P. Karst.) S.M. Berch (1983)
*Glomus viscosum*	Although this fungus has ostensibly been placed next to G. claroideum from sequence data, it seems the culture from which the evidence came also contained spores of other AM fungi. Consequently, verification should be waited for before placing it in any new genus.	T.H. Nicolson (1995)
*Glomus warcupii*		McGee (1986)
*Glomus zaozhuangianus*		F.Y. Wang & R.J. Liu (2002) [as 'zaozhuangianus']
*Glomus australe*	Basionym: Endogone australis Berk. (1859) [1860] Was synonymised with G. macrocarpum by Gerdemann & Trappe (1974). Because of lacking molecular evidence treated here as Glomus sensu lato of uncertain phylogenetic position.	(Berk.) S.M. Berch (1983)
*Otospora*	back to top	J. Palenzuela, N. Ferrol & Oehl (2008)	Diversisporaceae	Diversisporales
*Otospora bareae*	Unclear phylogenetic affiliation though placed among Diversispora based on two non-overlapping short SSU sequences. The type is un-interpretable due to parasitism and other degradation, and as yet there is no independent evidence is available to support its phylogenetic position; needs reinvestigation.	J. Palenzuela, N. Ferrol & Oehl (2008) [as bareai]	Diversisporaceae	Diversisporales
*Pacispora*	back to top	Oehl & Sieverd. (2004)	Pacisporaceae	Diversisporales
*Pacispora* *chimonobambusae*	Basionym: Glomus chimonobambusae C.G. Wu & Y.S. Liu (1995) Synonyms: Gerdemannia chimonobambusae C. Walker, Blaszk., Schuessler & Schwarzott (2004)	(C.G. Wu & Y.S. Liu) C. Walker, Vestberg & Schuessler (2007)	Pacisporaceae	Diversisporales
*Pacispora* *boliviana*		Oehl & Sieverd. (2004)	Pacisporaceae	Diversisporales
*Pacispora* *coralloidae*		Oehl & Sieverd. (2004)	Pacisporaceae	Diversisporales
*Pacispora* *franciscana*		Oehl & Sieverd. (2004)	Pacisporaceae	Diversisporales
*Pacispora* *patagonica*	Basionym: Glomus patagonicum Novas & Fracchia (2005) (link to publisher!)	(Novas & Fracchia) C. Walker, Vestberg & Schuessler (2007)	Pacisporaceae	Diversisporales
*Pacispora* *robigina*		Oehl & Sieverd. (2004)	Pacisporaceae	Diversisporales
*Pacispora* *scintillans*	Basionym: Glomus scintillans S.L. Rose & Trappe (1980) Synonyms: P. dominikii Oehl & Sieverd. (2004), Gerdemannia scintillans Walker, Błaszk., Schuessler & Schwarzott (2004), Glomus dominikii Błaszk. (1988)	(S.L. Rose & Trappe) C. Walker, Vestberg & Schuessler (2007)	Pacisporaceae	Diversisporales
*Paraglomus*	back to top	J.B. Morton & D. Redecker (2001)	Paraglomeraceae	Paraglomerales
*Paraglomus brasilianum*	Basionym: Glomus brasilianum Spain & J. Miranda (1996)	(Spain & J. Miranda) J.B. Morton & D. Redecker (2001)	Paraglomeraceae	Paraglomerales
*Paraglomus occultum* (complete paper here)	Basionym: Glomus occultum C. Walker (1982)	(C. Walker) J.B. Morton & D. Redecker (2001)	Paraglomeraceae	Paraglomerales
*Paraglomus laccatum* (link to commercial publisher not giving copyright clearance)	Basionym: Glomus laccatum Błaszk. (1988)	(Błaszk.) C. Renker, Błaszk. & F. Buscot (2007)	Paraglomeraceae	Paraglomerales
*Racocetra*	back to top	Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
*Racocetra* *castanea*	Basionym: Scutellospora castanea C. Walker (1993)	(C. Walker) Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
*Racocetra* *coralloidea*	Basionym: Gigaspora coralloidea Trappe, Gerd. & I. Ho (1974) Synonyms: Scutellospora coralloidea C. Walker & F.E. Sanders (1986)	(Trappe, Gerd. & I. Ho) Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
*Racocetra* *fulgida*	Basionym: Scutellospora fulgida Koske & C. Walker (1986)	(Koske & C. Walker) Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
*Racocetra* *gregaria*	Basionym: Gigaspora gregaria N.C. Schenck & T.H. Nicolson (1979) Synonyms: Scutellospora gregaria C. Walker & F.E. Sanders (1986)	(N.C. Schenck & T.H. Nicolson) Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
*Racocetra* *persica*	Basionym: Gigaspora persica Koske & C. Walker (1985) Synonyms: Scutellospora persica C. Walker & F.E. Sanders (1986)	(Koske & C. Walker) Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
Racocetra tropicana		Oehl, B.T.Goto & G.A.Silva (2011)	Gigasporaceae	Diversisporales
*Racocetra* *verrucosa*	Basionym: Gigaspora verrucosa Koske & C. Walker (1985) Synonyms: Scutellospora verrucosa C. Walker & F.E. Sanders (1986)	(Koske & C. Walker) Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
*Racocetra* *weresubiae*	Basionym: Scutellospora weresubiae Koske & C. Walker (1986) Not established in culture, but specimens collected from sand dunes in Argentina were clearly of this species. Msiska & Morton (2010) removed this species from Racocetra, but Schüßler and Walker (2010) confirmed it to be in a monophyletic clade with Racocetra, therefore it was places back in that genus.	(Koske & C. Walker) Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
*Racocetra*	species of uncertain position placed in the genus on the grounds of morphological characters that have been criticised by Msiska & Morton (2010).		Gigasporaceae	Diversisporales
*Racocetra* *alborosea*	Basionym: Gigaspora alborosea Ferrer & R.A. Herrera (1981) Synonyms: Scutellospora alborosea C. Walker & F.E. Sanders (1986)	(Ferrer & R.A. Herrera) Oehl, F.A. Souza & Sieverd. (2008)	Gigasporaceae	Diversisporales
*Racocetra beninensis*		Oehl, Tchabi & Lawouin (2009)
*Racocetra intraornata*